EQUATIONS TO CALCULATE PHOTOSYNTHESIS



Transpiration.-

	          Fd·e             K                  Fd       de
                 ------- + [----------------] [Vt -  ---- Vg] ----
                  100·P     8.314 (Ta + 273)          Fx       dt 
	    E = ------------------------------------------------------
                                          e
                                  S [1 - --- ]
                            	          P                                                                 

	Where:
		Ta= Chamber air temperature (°C).
		Fd= Flow through desiccant µmol/s.
	 	 e= Vapor pressure of air (mb).    
             de/dt= Rate of change of e.
                 E= Transpiration rate (mol/m²/s).
		 P= Atmospheric pressure (mb).
		 S= Leaf area (cm²).
                Vt= Total volume (cm3).
                Fx= Max flow rate (µmol/s).   
                Vg= IRGA volume (cm3).
                 K= Absorption coefficient.


Stomatal Conductance.-

	The equation to determine the stomatal conductance to water vapor
takes into account the convective effect at the stomatal pores caused by 
the transpiration.


                               1
               gsw =  ---------------------
                            A  -  B



	                    es(Tl)  -  e         
                A  =   -------------------------
	                          es(Tl) + e    
		        E [P  -  ------------ ]               	  	 	                             2                                                                                    
	                               2


	                  (K² + 1)
                B  =  ------------------
	                gbwo (K + 1)²
		                     	  	 	                             2                                                                                    


Where:
		gsw in mol m²/s 
		 Tl= Leaf temperature (°C).
		  e= Vapor pressure in chamber (mb).
		 es= Saturation vapor pressure function (mb)
	       gbwo= One side boundary layer conductance (mol/m²/s). 
                  K= Stomatal ratio.


Photosynthesis.-
	
	In this case the term (dc/dt) is included to simulate transient conditions in a closed system. 
		
             Fd · e · C             P· G · Vt           dc
      A =  --------------  -  --------------------  -  ----  -  C·E·G
            100 · S · P        8.314 (Ta + 273) S       dt


	Where:
		 A in µmol/m²/s
		C= CO2 concentration (µl/l)
		


Intercellular CO2 Content.-

                                         E
                               [ gtc -  ---  Ca - A ]
                                         2
			 Ci = -------------------------
                                           E    
      		                  [ gtc + --- ]      
         	       		           2

	Where:
	      gtc= Total conductance to CO2

	The total conductance to CO2 is calculated from the conductances for water vapor:

		         1     1.6     1.37
                        --- = ----- + ------
         	        gtc    gsw     gbw

	Where:
		 1.37= Ratio of diffusivities of CO2 and water vapor in 
                       the boundary layer.
		  1.6= Ratio of diffusivities of CO2 and water vapor in air.
					

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